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Dogs tilt their head to better determine the vertical location of a sound.

That's always been my hypothesis.

Dogs have two ears, spaced horizontally apart such that triangulation can be used to determine the origin of the sound by exploiting the small time difference in time for the sound to reach both ears.

But vertically speaking the ears are in the same position. So to be able to better judge the vertical position of the origin of the sound, they title their heads in order to place the ears apart on the vertical axis.




That seems to accord with what the fox does at 1:08 here: https://youtu.be/D2SoGHFM18I




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